New York Metropolitan Flora

Nomenclature

Fraxinus L., Sp. Pl. 1057. 1753. Gen Pl., ed. 5, 477. 1754. LECTOTYPE: Fraxinus excelsior L. designated by Britton & Brown (1913)

Calycomelia Kostel., Allg. Med.-Pharm. Fl. 3: 1003. 1834. TYPE: Not designated.

Leptalix Raf., New Fl. 3: 93. 1838 (“1836”). TYPE: Not designated.

Aplilia Raf., New Fl. 3: 93. 1838 (“1836”). TYPE: Not designated.

Key to the species of Fraxinus

1. Leaflets sessile...2
1. Leaflets with a petiole...3

2. Rachis at base of leaflets with thick reddish tomentum; flowers dioecious...Fraxinus nigra
2. Rachis without conspicuous tomentum; flower polygamous...Fraxinus excelsior

3. Leaf scar upper margin sunken with bud down into scar...Fraxinus americana
3. Leaf scar upper margin straight, with bud above scar...4

4. Current year’s twigs glabrous or with white or gray tomentum...Fraxinus pennsylvanica
4. Current year’s twigs with brownish-yellow tomentum...Fraxinus profunda

List of Fraxinus Species

References to Fraxinus

• Abrams, M. D.; Kubiske, M. E.; Mostoller, S. A. 1994. Relating wet and dry year ecophysiology to leaf structure in contrasting temperate tree species. Ecology 75: 123-33.
• Abrams, M. D.; Kubiske, M. E.; Steiner, K. C. 1990. Drought adaptations and responses in five genotypes of Fraxinus pennsylvanica Marsh.: Photosythesis, water relations and leaf morphology. Tree Physiology 6: 305-15.
• Ahlgren, C. E. 1957. Phenological observations of nineteen native tree species in northeastern Minnesota. Ecology 38: 622-8.
• Alexander, N. L.; Flint, H. L.; Hammer, P. A. 1984. Variation in cold-hardiness of Fraxinus americana stem tissue according to geographic origin. Ecology 65: 1087-92.
• Altpeter, L. S. 1944. Use of vegetation in control of streambank erosion in Northern New England. J. Forest. 42(2): 99-107.
• Andersen, C. P.; Sucoff, E. I.; Dixon, R. K. 1987. The influence of low soil temperatures on the growth of vescular-arbuscular mycorrhizal Fraxinus pennsylvanica. Canad. J. Forest Res. 17(8): 951-6.
• Anderson, E.; Whelden, C. M. 1936. Studies in the genus Fraxinus. II. Data on the flowering and fruiting habits of three American species of ash- of possible economic importance in the production of rapid-growing forest trees. J. Heredity 27: 473-4.
• Armstrong, J. E. 1982. Polyploidy and wood anatomy of mature white ash, Fraxinus americana. Wood & Fiber 14(4): 331-9.
• Balter, H.; Loeb, R. E. 1983. Arboreal relationships on limestone and gneiss in northern New Jersey and southeastern New York. Bull. Torrey Bot. Club 110: 370-9.
• Baltzer, J. L.; Thomas, S. C. 2005. Leaf optical responses to light and soil nutrient availability in temperate deciduous trees. Amer. J. Bot. 92: 214-223.
• Bartlett, G. A. 1997. Crown development and architecture of Fraxinus pennsylvanica var. subintegerrima (Vahl) Fern. At different stages of growth and under different environmental conditions. M.Sc. Thesis Univ. Manitoba, Winnipeg,
• Bartlett, G. A.; Remphrey, W. R. 1998. The effect of reduced quantities of photosynthetically active radiation on Fraxinus pennsylvanica growth and architecture. Canad. J. Bot. 76: 1359-1365.
• Barton, L. V. 1945. Viability of seeds of Fraxinus after storage. Contr. Boyce Thompson Inst. Pl. Res. 13: 427-32.
• Bazzaz, F. A.; Miao, S. L. 1993. Successional status, seed size, and responses of tree seedlings to CO2, light, and nutrients. Ecology 74: 104-12.
• Bell, D. T.; Johnson, F. L. 1975. Phenological patterns in the trees of the streamside forest. Bull. Torrey Bot. Club 102: 187-93.
• Benedict, L.; David, R. 2003. Propagation protocol for black ash, Fraxinus nigra Marsh. Native Plants J. 4: 100-103.
• Berry, E. W. 1918. The history of linden and ash. Pl. World 21: 163-75.
• Bessey, C. E. 1914. Tricarpellary ash-fruits. American Botanist 20: 21.
• Black, C. L.; Beckmann, R. L. 1983. The variability of nuclear DNA and its implications for polyploidy in white ash (Fraxinus americana L.: Oleaceae). Amer. J. Bot. 70: 1420-3.
• Black-Schaefer, C. L.; Beckmann, R. L. 1989. Foliar flavonoids and the determination of ploidy and gender in Fraxinus americana and F. pennsylvanica (Oleaceae). Castanea 54: 115-8.
• Boerner, R. E. J.; Brinkman, J. A. 1996. Ten years of tree seedling establishment and mortality in an Ohio deciduous forest complex. Bull. Torrey Bot. Club 123: 309-17.
• Bowden, W. M. 1940. Diploidy, polyploidy, and winter hardiness relationships in the flowering plants. Amer. J. Bot. 27: 357-71.
• Britton, Nathaniel L. 1892. The ash trees of Staten Island. Proc. Nat. Sci. Assoc. Staten Island 3: 21-2.
• Bruederle, L. P.; Stearns, F. W. 1985. Ice storm damage to a southern Wisconsin mesic forest. Bull. Torrey Bot. Club 112(2): 167-75.
• Bunce, J. A.; Miller, L. N. 1976. Differential effects of water stress on respiration in the light in woody plants from wet and dry habitats. Canad. J. Bot. 54(21): 2457-64.
• Bunce, J. A.; Miller, L. N.; Chabot, B. F. 1977. Competative exploitation of soil water by five eastern North American tree species. Bot. Gaz. 138(2): 168-73.
• Burton, P. J.; Bazzaz, F. A. 1991. Tree seedling emergence on interactive temperature and moisture gradients and in patches of old-field vegetation. Amer. J. Bot. 78(1): 131-49.
• Buton, P. J.; Bazzaz, F. A. 1995. Ecophysiological responses of tree seedlings invading different patches of old-field vegetation. J. Ecol. 83: 99-112.
• Canham, C. D.; McAninch, J. B.; Wood, D. M. 1994. Effects of the frequency, timing, and intensity of simulated browsing on growth and mortality of tree seedlings. Canad. J. Forest Res. 24: 817-25.
• Carter, K. K. 1996. Provenance tests as indicators of growth response to climate change in 10 north temperate tree species. Canad. J. Forest Res. 26: 1089-95. (French summary)
• Castello, J. D.; Silverborg, S. B.; Manion, P. D. 1985. Intensification of ash decline in New York State from 1962 through 1980. Plant Dis. 69: 243-6.
• Chadwick, L. C. 1939. New England hurricane damage to trees. Arborist's News 4(1): 1-4.
• Chappelka, A. H.; Chevone, B. I. 1986. White ash seedling growth response to ozone and simulated acid rain. Canad. J. Forest Res. 16: 786-90.
• Choat, B. et.al. 2006. Direct measurements of intervessel pit membrane hydraulic resistance in two angiosperm tree species. Amer. J. Bot. 93: 993-1000.
• Christianson, J. D. 1971. The effect of surface fires on three species of hardwood tree seedlings. William L. Hutcheson Memorial For. Bull. 2: 1-5.
• Clausen, K. E. 1984. Survival and early growth of white ash provenances and progenies in 19 plantations. Canad. J. Forest Res. 14: 775-82.
• Clausen, K. E.; Kung, F. H.; Bey, C. F.; Daniels, R. A. 1981. Variation in white ash. Silvae Genet. 30: 93-7.
• Cluzeau, C.; Le Goff, N.; Ottorini, J. M. 1994. Development of primary branches and crown profile of Fraxinus excelsior. Canad. J. Forest Res. 24(12): 2315-23.
• Coladoanto, M. 1994. Fraxinus nigra. ()
• Cook, D. B. 1941. The period of growth in some northeastern trees. J. Forest. 39: 956-959.
• Curtis, R. O.; Rushmore, F. M. 1958. Some effects of stand density and deer browsing on reproduction in an Adirondack hardwood stand. J. Forest. 56: 116-21.
• Davidson, C. G.; Remphrey, W. R. 1990. An analysis of architectural parameters of male and female Fraxinus pennsylvanica in relation to crown shape and crown location. Canad. J. Bot. 68: 2035-43. (French summary)
• Davies, W. J.; Kozlowski, T. T. 1974. Stomatal responses of five woody angiosperms to light intensity and humidity. Canad. J. Bot. 52(7): 1525-34.
• Davison, S. E. 1981. Tree seedling survivorship at Hutcheson Memorial Forest New Jersey. William L. Hutcheson Memorial For. Bull. 6: 4-7.
• Dayton, W. A. 1954. Some more notes on United States ashes (Fraxinus). J. Wash. Acad. Sci. 44: 385-90.
• De Jong, P. C. 1990. The genus Fraxinus. Dendroflora 26: 30-9. (In Dutch)
• De Steven, D. 1991. Experiments on mechanisms of tree establishment in old-field succession: seedling emergence. Ecology 72: 1066-75.
• De Steven, D. 1991. Experiments on mechanisms of tree establishment in old-field succession: seedling survival and growth. Ecology 72: 1076-88.
• Delucia, E. H.; Sipe, T. W.; Herrick, J.; Maherali, H. 1998. Sapling biomass allocation and growth in the understory of a deciduous hardwood forest. Amer. J. Bot. 85: 955-963.
• Den Uyl, D. 1962. Survival and growth of hardwood plantations on strip mine spoil banks of Indiana. J. Forest. 60(9): 603-6.
• Denneler, B; Bergeron, Y.; Begin, Y. 1999. An attempt to explain the distribution of the tree species composing the riparian forests of Lake Duparquet, southern boreal region of Quebec, Canada. Canad. J. Bot. 77: 1744-1755.
• Dolan, R. W. 1994. Effects of proscribed burn on tree- and herb-layer vegetation in a post oak (Quercus stellata) dominated flatwoods. Proc. Indiana Acad. Sci. 103: 25-32.
• Dolia, N. I. 1953. In regard to the physiology of germination of ash (Fraxinus) seed embryos. Dokl. Akad. Nauk SSSR 88: 729-32. (In Russian)
• Echternach, J. L.; Rose, R. K. 1987. Use of woody vegetation by beavers in southeastern Virginia USA. Virginia J. Sci. 38: 226-232.
• Facey, V. 1950. Abscission of leaves in Fraxinus americana L. New Phyt. 49: 103-16.
• Fernald, M. L.; Schubert, B. G. 1948. Studies of American types in British herbaria. Rhodora 50: 149-76, 181-208, 217-33.
• Fitzgerald, C. H.; Reines, M. 1969. A comparative study of the flavonoid content of Fraxinus americana and Fraxinus pennsylvanica. Castanea 34: 192-4.
• Flower, F. B.; Gilman, E. F.; Leone, I. A. 1981. Landfill gas, what it does to trees and how its injurious effects may be prevented. J. Arboric. 7(2): 43-52.
• Gardescu, S.; Marks, P. L. 2004. Colonization of old fields by trees vs. shrubs: seed dispersal and seedling establishment. J. Torrey Bot. Soc. 131: 53-68.
• Gardner, G. 1977. The reproductive capacity of Fraxinus excelsior on the Derbyshire limestone. J. Ecol. 65: 107-18.
• Godman, R. M.; Mattson, G. A. 1970. Periodic growth of hardwood influenced by cold weather. J. Forest. 68(2): 86-7.
• Goldsmith, F. B.; Boudreau, P. 1979. Height growth and apical damage of white ash (Fraxinus americana L.) from various latitudes outplanted in New Brunswick. Canad. J. Forest Res. 9: 27-30.
• Green, D. S. 1980. The terminal velocity and dispersal of spinning samaras. Amer. J. Bot. 67: 1218-24.
• Greene, D. F.; Johnson, E. A. 1992. Fruit abscission in Acer saccharinum with reference to seed dispersal. Canad. J. Bot. 70(11): 2277-83.
• Greene, D. F.; Johnson, E. A. 1990. The dispersal of winged fruits and seeds differing in autorotative behavior. Canad. J. Bot. 68: 2693-7. (French summary)
• Griffith, R. S. 1991. Fraxinus americana. ()
• Guenther, K. W. 1951. An investigation of the tolerance of white ash reproduction. J. Forest. 49: 576-7.
• Harborne, J. B. 1980. A chemotaxonomic survey of flavonoids in leaves of the Oleaceae. J. Linn. Soc. Bot. 81: 155-67.
• Hardin, J. W.; Beckmann, R. L. 1982. Atlas of foliar surface features in woody plants, V. Fraxinus (Oleaceae) of eastern North America. Brittonia 34: 129-40.
• Harris, J. R.; Bassuk, N. L.; Zobel, R. W.; Whitlow, T. H. 1995. Root and shoot growth periodicity of Green Ash, Scarlet Oak, Turkish Hazelnut, and Tree Lilac. J. Amer. Soc. Hort. Sci. 120: 211-6.
• Heaslip, M. B. 1959. Effects of seed irradiation on germination and seedling growth of certain deciduous trees. Ecology 40(3): 383-8.
• Hemberg, T. 1949. Growth-inhibiting substances in terminal buds of Fraxinus. Physiol. Pl. (Copenhagen) 2: 37-44.
• Hengst, G. E.; Dawson, J. O. 1994. Bark properties and fire resistance of selected tree species from the central hardwood region of North America. Canad. J. Forest Res. 24: 688-96.
• Hildebrand, E.; Skelly, J. M.; Fredericksen, T. S. 1996. Foliar response of ozone-sensitve hardwood tree species from 1991 to 1993 in the Shenandoah National Park, Virginia. Canad. J. Forest Res. 26: 658-69. (French summary)
• Hoppes, W. G. 1988. Seedfall pattern of several species of bird-dispersed plants in an Illinois woodland. Ecology 69: 320-329.
• Hornbeck, J. W.; Smith, R. B.; Federer, C. A. 1988. Growth trends in 10 species of trees in New England, 1950-1980. Canad. J. Forest Res. 18: 1337-40.
• Hosner, J. F. 1958. The effects of complete inundation upon seedlings of six bottomland tree species. Ecology 39: 371-3.
• Hosner, J. F. 1959. Survival, root, and shoot growth of six bottomland tree species following flooding. J. Forest. 57: 927-8.
• Hough, A. F.; Forbes, R. D. 1943. The ecology and silvics of forests in the high plateaus of Pennsylvania. Ecol. Monogr. 13: 299-320.
• Houston, D. R. 1985. Dieback and declines of urban trees. J. Arboric. 11(3): 65-72.
• Jacobs, K.; Danielson, D. 2002. Variation in field susceptibility of native and exotic ash species to anthracnose. J. Arboric. 28: 35-40.
• Jeandroz, S.; Roy, A.; Bousquet, J. 1997. Phylogeny and phylogeography of the circumpolar genus Fraxinus (Oleaceae) based on internal transcribed spacer sequences of nuclear ribosomal DNA. Molec. Phylogenet. Evol. 7: 241-51.
• Johnson, F. L.; Bell, D. T. 1976. Tree growth and mortality in the streamside forest. Castanea 41: 34-41.
• Johnson, W. C. 1988. Estimating dispersibility of Acer, Fraxinus and Tilia in fragmented landscapes from patterns of seedling establishment. Landscape Ecol. 1(3): 175-88.
• Jurik, T. W.; Weber, J. A.; Gates, D. M. 1988. Effects of temperature and light on photosynthesis of dominant species of a northern hardwood forest. Bot. Gaz. 149: 203-8.
• Karnosky, D. F. 1981. Chamber and field evaluations of air pollution tolerances of urban trees. J. Arboric. 7: 99-105. (Sulphur dioxide, ozone, & oxidants)
• Karnosky, D. F.; Steiner, K. C. 1981. Provenance and family variation in response of Fraxinus americana and Fraxinus pennsylvanica to ozone and sulfur dioxide. Phytopathology 71: 804-7.
• Keeley, J. E.; Pallardy, S. G. 1979. Stomatal responses of Fraxinus pennsylvanica seedlings during and after flooding. Physiol. Pl. (Copenhagen) 46: 155-158.
• Kennedy, G.; Bergeron, S. 1991. Tree rings as monitors of heavy metal air pollution histories. Journal of Radioanalytical and Nuclear Chemistry 151: 337-344.
• Kienholz, R. 1941. Seasonal course of height growth in some hardwoods in Connecticut. Ecology 22: 249-58.
• Killingbeck, K. T. 1985. Autumnal resorption and accretion of trace metals in gallery forest tress. Ecology 66: 283-6. (Cu, Fe, Mn, & Zn)
• King, J. L. 1994. Persistence of black cherry, white ash, and tulip poplar in an old-growth beech-maple forest. , Thesis Univ. Illinois, Chicago,
• Kozlowski, T. T.; Pallardy, S. G. 1979. Effects of low temperature on leaf diffusion resistance of Ulmus americana and Fraxinus pennsylvanica seedlings. Canad. J. Bot. 57: 2466-70.
• Kramer, P. J. 1943. Amount and duration of growth of various species of tree seedlings. Pl. Physiol. (Lancaster) 18: 239-51.
• Kung, F. H.; Clausen, K. E. 1983. Provenance x environment interactions in white ash.
• Lanier, G. N.; Schubert, D. C.; Manion, P. D. 1988. Dutch elm disease and elm yellows in central New York. Plant Dis. 72: 189-94.
• Larson, P. R. 1985. Rachis vascularization and leaflet venation in developing leaves of Fraxinus pennsylvaica. Canad. J. Bot. 63: 2383-92.
• Lawrey, J. D. 1977. Trace metal accumulation by plant species from a coal strip-mining area in Ohio. Bull. Torrey Bot. Club 104: 368-375.
• Lemon, P. C. 1961. Forest ecology of ice storms. Bull. Torrey Bot. Club 88(1): 21-9.
• Leone, I. A.; Flower, F. B.; Arthur, J. J.; Gilman, E. F. 1977. Damage to woody species by anaerobic landfill gases. J. Arboric. 3(12): 221-5.
• Leverett, R. 1996. The white ash. Wild Earth 6(2): 37-42.
• Lingelsheim, A. 1907. Vorarbeiten zu einer Monographie der Gattung Fraxinus. Bot. Jahrb. Syst. 40: 185-223. (In German)
• Little, E. L. 1952. Notes on Fraxinus (ash) in the United States. J. Wash. Acad. Sci. 42: 369-80.
• Loats, K. V.; Rebbeck, J. 1999. Interactive effects of ozone and elevated carbon dioxide on the growth and physiology of black cherry, green ash, and yellow poplar seedlings. Environmental Pollution 106: 237-248.
• Logan, K. T. 1973. Growth of tree seedlings as affected by light intensity V. White ash, beech, eastern hemlock, and general conclusions.
• Luken, J. O.; Hinton, A. C.; Baker, D. G. 1992. Response of woody plant communities in power-line corridors to frequent anthropogenic disturbance. Ecological Applications 2: 356-62.
• Marks, P. L. 1975. On the relation between extension growth and successional status of deciduous trees of the northeastern United States. Bull. Torrey Bot. Club 102: 172-7.
• Marshall, P. E.; Kozlowski, T. T. 1977. Changes in structure and function of epigeous cotyledons of woody angiosperms during early seedling growth. Canad. J. Bot. 55(2): 208-15.
• Marshall, P. E.; Kozlowski, T. T. 1976. Compositional changes in cotyledons of woody angiosperms. Canad. J. Bot. 54(21): 2473-7.
• Marshall, P. E.; Kozlowski, T. T. 1975. Changes in mineral contents of cotyledons and young seedlings of woody angiosperms. Canad. J. Bot. 53(18): 2026-31.
• Matlack, G. R. 1987. Diaspore size, shape, and fall behavior in wind-dispersed plant species. Amer. J. Bot. 74(8): 1150-60.
• Matteoni, J. A.; Sinclair, W. A. 1985. Role of the mycoplasmal disease, ash yellows, in decline of white ash in New York State. Phytopathology 75: 355-60.
• McApline, R. G. 1961. Yellow-poplar seedlings intolerant to flooding J. Forest. 59(8): 566-8. (Also Fraxinus & Liquidambar)
• McClendon, J. H.; McMillen, G. G. 1982. The control of leaf morphology and the tolerance of shade by woody plants. Bot. Gaz. 143(1): 79-83.
• McCormac, J. S.; Bissell, J. K.; Stine, S. J. 1995. The status of Fraxinus tomentosa (Oleaceae) in Ohio with notes on its occurrence in Michigan and Pennsylvania. Castanea 60: 70-8.
• McCormick, L. H.; Bowersox, T. W. 1997. Grass or fern competition reduce growth and survival of planted tree seedlings.
• McCutchen, C. W. 1977. The spinning rotation of ash and tulip-tree samaras. Science 197: 691-2.
• McDonnell, M. J.; Stiles, E. W. 1983. The structural complexity of old field vegetation and the recruitment of bird-dispersed plant species. Oecologia 56: 109-16.
• McIninch, S. M.; Garbisch, E. W.; Biggs, D. 1994. The benefits of wet-acclimating woody wetland plant species. Wetland Journal 6: 19-23.
• Meiners, S. J.; Gorchov, D. L. 1998. Effects of distance to Juniperus virginiana on the establishment of Fraxinus and Acer seedlings in old fields. Amer. Midl. Naturalist 139: 353-64.
• Meiners, S. J.; Stiles, E. W. 1997. Selective predation on the seeds of woody plants. J. Torrey Bot. Soc. 124: 67-70.
• Merrill, E. K. 1986. Heteroblastic seedlings of green ash. I, II, & III. Canad. J. Bot. 64: 2645-68.
• Merrill, E. K. 1990. Structure and development of terminal bus scales in green ash. Canad. J. Bot. 68: 12-20. (French summary)
• Miller, G. N. 1955. The genus Fraxinus, the ashes, in North America, north of Mexico. Cornell Univ. Agr. Exp. Sta. Mem. 335. Cornell Univ., Ithaca, NY. , 64 pages.
• Miller, G. N. 1953. A taxonomic revision of the genus Fraxinus in North America north of Mexico. Ph.D. Dissertation Cornell Univ., Ithaca, NY200 p.
• Myster, R. W. 1993. Effects of litter, distance, density and vegetation patch type on postdispersal tree seed predation in old fields. Oikos 66: 381-388.
• Nesom, G. L. 2010. Fraxinus biltmoreana and Fraxinus smallii (Oleaceae), forest trees of the eastern United States. Phytoneuron 2010-51: 1-30.
• Nesom, G. L. 2010. Notes on Fraxinus profunda (Oleaeceae). Phytoneuron 2010-32: 1-6.
• Nikolaev, E. V. 1982. Morphogenesis and evolution of the genus Fraxinus (Oleaceae). Bot. Zhurn. (Moscow & Lenengrad) 67(4): 419-32. (In Russian; English summary)
• Nikolaeva, M. G. 1951. The causes of seed dormancy in Acer negundo L., Fraxinus pensylvanica Marsh., and Berberis vulgaris f. atropurpurea Regel. Trudy Bot. Inst. Akad. Nauk SSSR, Ser. 4, Eksper. Bot. 8: 234-56.
• Nilolaeva, M. G. 1958. The biology of germination of Fraxinus seeds and its bearing on the taxonomic position and the distribution of species of this genus. Bot. Zhurn. (Moscow & Lenengrad) 43: 679-83. (In Russian)
• Ostfeld, R. S.; Manson, R. H.; Canham, C. D. 1997. Effects of rodents on survival of tree seeds and seedlings invading old fields. Ecology 78: 1531-42.
• Paratley, R. D. 1986. Vegetation-environment relations in a conifer swamp in central New York. J. Torrey Bot. Soc. 113: 357-371.
• Parker, J. 1968. Drought resistance of roots of white ash, sugar maple, and red oak.
• Pickett, S. T. A.; Kempf, J. S. 1980. Branching patterns in forest shrubs and understory trees in relation to habitat. New Phyt. 86: 219-228.
• Pope, P. E.; Chaney, W. R.; Rhodes, J. D.; Woodhead, S. H. 1983. The mycorrhizal dependency of four hardwood tree species. Canad. J. Bot. 61: 412-7.
• Prusinkiewicz, P. W.; Remphrey, W. R.; Davidson, C. G.; Hammel, M. S. 1994. Modeling an architecture of expanding Fraxinus pennsylvanica shoots using L-systems. Canad. J. Bot. 72(5): 701-14.
• Rankin, W. T.; Pickett, S. T. A. 1989. Time of establishment of red maple (Acer rubrum) in early oldfield succession. Bull. Torrey Bot. Club 116: 182-6. (Also Carya, Juniperus, Nyssa, Prunus, & Quercus)
• Remphrey, W. R. 1989. Shoot ontogeny in Fraxinus pennsylvanica (green ash). II. Development of the inflorescence. Canad. J. Bot. 67: 1966-78. (French summary)
• Remphrey, W. R. 1989. Shoot ontogeny in Fraxinus pennsylvanica (green ash). I. Seasonal cycle of terminal meristem activty. Canad. J. Bot. 67: 1624-32. (French summary)
• Remphrey, W. R.; Bartlett, G. A.; Davidson, C. G. 2002. Shoot morphology and fate of buds in relation to crown location in young Fraxinus pennsylvanica var. subintegerrima. Canad. J. Bot. 80: 1274-1282.
• Remphrey, W. R.; Davidson, C. G. 1992. Branch architecture and its relation to shoot-tip abortion in mature Fraxinus pennsylvanica. Canad. J. Bot. 70(6): 1147-53.
• Remphrey, W. R.; Davidson, C. G. 1994. Shoot and leaf growth in Fraxinus pennsylvanica and its relation to crown location and pruning. Canad. J. Forest Res. 24: 1997-2005.
• Remphrey, W. R.; Davidson, C. G.; Blouw, M. J. 1987. A classification and analysis of crown form in green ash (Fraxinus pennsylvanica). Canad. J. Bot. 65: 2188-95.
• Rhoads, A. F. 1976. Forest species show a delayed response to cement dust in the soil. J. Arboric. 2(10): 197-9.
• Ricklefs, R. E.; Matthew, K. K. 1982. Chemical characteristics of the foliage of some deciduous trees in southeastern Ontario. Canad. J. Bot. 60: 2037-45.
• Roberds, J. H.; Hyun, J. O.; Namkoong, G.; Rink, G. 1990. Height response functions for white ash provenances grown at different latitudes. Silvae Genet. 39: 121-9.
• Roberts, B. R. 1980. Trees as biological filters. J. Arboric. 6(1): 20-3. (Sulphur dioxide & ozone)
• Rohwer, J. G. 1993. A preliminary survey of the fruits and seed of the Oleaceae. Bot. Jahrb. Syst. 115(2): 271-91.
• Rosario, L. C. 1988. Fraxinus pennsylvanica. ()
• Rudolf, P. O. 1929. A study of the seedling root systems of certain forest trees. M.S. Thesis Cornell Univ., Ithaca, NY60 p.
• Santamour, F. S. 1962. The relation between polyploidy and morphology in white and biltmore ashes. Bull. Torrey Bot. Club 89: 228-32.
• Santamour, F. S. 1963. Thirteen-year growth of some green ash provenances in the Northeast.
• Sargent, C. S. 1894. Fraxinus. Silva N. Am. 6: 25-58.
• Sax, K.; Abbe, E. C. 1932. Chromosome numbers and the anatomy of the secondary xylem in the Oleaceae. J. Arnold Arbor. 13: 37-48.
• Schrauwen, J. A. M.; Sondheimer, E. 1975. The development of organelles in embryos of Fraxinus americana. Zeitschr. Pflanzenphysiol. 75: 67-77.
• Sena Gomes, A. R.; Kozlowski, T. T. 1980. Growth responses and adaptations of Fraxinus pennsylvanica seedlings to flooding. Pl. Physiol. (Lancaster) 66: 267-71.
• Shanklin, J.; Kozlowski, T. T. 1984. Effect of temperature preconditioning on response of Fraxinus pennsylvanica seedlings to SO2. Environm. Pollut., A. 36: 311-26.
• Shumway, D. L.; Steiner, K. C.; Abrams, M. D. 1991. Effects of drought stress on hydraulic architecture of seedlings from five populations of green ash. Canad. J. Bot. 69(10): 2158-64.
• Silverborg, S. B.; Ross, E. W. 1968. Ash dieback disease development in New York State. Pl. Dis. Reporter 52: 105-7.
• Sinclair, W. A. 1964. Comparisons of recent declines of white ash, oaks and sugar maple in northeastern woodlands. Cornell Plantations 20: 62-7.
• Sinclair, W. A.; Griffiths, H. M.; Lee, I. M. 1994. Mycoplasmlike organisims as causes of slow growth and decline of trees and shrubs. J. Arboric. 20(3): 176-89. (Fraxinus & Ulmus)
• Sondheimer, E.; Blank, G. E.; Galson, E. C.; Sheets, F. M. 1970. Metabolically active glucosides in Oleaceae seeds. 1. The effects of germination, growth, and hormone treatments. Pl. Physiol. (Lancaster) 45: 658-62.
• Steen, D. A. et al. et.al. 1995. The big trees of Michigan: 11. Fraxinus pennsylvanica L. Michigan Bot. 34: 144-6.
• Steinbauer, G. P. 1943. Frequency of polyembryony in Fraxinus seeds. Bot. Gaz. 105: 285.
• Steinbauer, G. P. 1937. Dormancy and germination of Fraxinus seeds. Pl. Physiol. (Lancaster) 12: 813-24.
• Steiner, K. C. et al. et.al. 1988. Juvenile performance in a range-wide provenance test of Fraxinus pennsylvanica Marsh. Silvae Genet. 37: 104-11.
• Sterrett, W. D. 1915. The ashes; their characteristics and management.
• Tardif, J.; Bergeron, Y. 1993. Radial growth of Fraxinus nigra in a Canadian boreal floodplain in response to climatic and hydrological fluctuations. J. Veg. Sci. 4: 751-8.
• Tardif, J.; Bergeron, Y. 1992. Ecological analysis of black ash populations (Fraxinus nigra) from the floodplains of Lake Duparquet in northwestern Quebec. Canad. J. Bot. 70: 2294-302.
• Tardif, J.; Dery, S.; Bergeron, Y. 1994. Sexual regeneration of black ash (Fraxinus nigra Marsh.) in a boreal floodplain. Amer. Midl. Naturalist 132: 124-35.
• Taylor, H. 1945. Cytotaxonomy and phylogeny of the Oleaceae. Brittonia 5: 337-67.
• Taylor, S. M. 1971. Ecological seperation and post-settlement habitat changes of Fraxinus americana and Fraxinus pennsylvanica in Washtenaw County, Michigan. Amer. J. Bot. 58(5): 466-7. (Abstract)
• Taylor, S. M. 1972. Ecological and genetic isolation of Fraxinus americana and Fraxinus pennsylvanica. Ph.D. Dissertation Univ. of Michigan, Ann Arbor, MI,
• Taylor, S. M. O. 1973. Ecological and genetic isolation of Fraxinus americana and Fraxinus pennsylvanica. Dissertation Abstracts Int., B 33(11): 5176. (Abstract)
• Tobiessen, P.; Buchsbaum, S. 1976. Ash dieback and drought. Canad. J. Bot. 54(6): 543-5.
• Tobiessen, P.; Kana, T. M. 1974. Drought-stress avoidance in three pioneer tree species. Ecology 55(3): 667-70.
• Tobiessen, P.; Werner, M. B. 1980. Hardwood seedling survival under plantations of scotch pine and red pine in central New York. Ecology 61(1): 25-9.
• Tryon, E. H.; True, R. P. 1968. Radial increment response of Appalachian hardwood species to a spring freeze. J. Forest. 66(6): 488-91.
• U. S. Forest Service. 1957. White ash (Fraxinus americana).
• Vaartaja, O. 1959. Evidence of photoperiodic ecotypes in trees. Ecol. Monogr. 29: 91-111.
• Vanstone, D. E. 1974. Seed dormancy in black ash (Fraxinus nigra). Proc. Ann. Meet. West Canad. Soc. Hort. 30: 77084.
• Vanstone, D. E.; DeLacroix, L. J. 1975. Embryo immaturity and dormancy of black ash. J. Amer. Soc. Hort. Sci. 100: 630-2.
• Vrecenak, A. J.; Vodak, M. C.; Fleming, L. E. 1989. The influence of site factors on the growth of urban trees. J. Arboric. 15(9): 206-9.
• Waldron, G.; Gartshore, M.; Colthurst, K. 1996. Pumpkin ash, Fraxinus profunda, in southwestern Ontario. Canad. Field-Naturalist 110: 615-9.
• Wallander, E. 2008. Systematics of Fraxinus (Oleaceae) and evolution of dioecy. Pl. Syst. Evol. 273: 25-49.
• Ward, J. S. 1997. White ash (Fraxinus americana L.) survival and growth in unmanaged upland forests.
• Wardle, P. 1959. The regeneration of Fraxinus excelsior in woods with a field layer of Mercurialis perennis. J. Ecol. 47: 483-97.
• Wells, O. O. 1986. Geographic variation in green ash in the southern coastal plain of the United States. Silvae Genet. 35: 165-9.
• Wenzig, T. 1883. Die Gattung Fraxninus Tourn. Bot. Jahrb. Syst. 4: 165-88. (In German)
• Werner, P. A.; Harbeck, A. L. 1982. The pattern of tree seedling establishment relative to staghorn sumac cover in Michigan old fields. Amer. Midl. Naturalist 108: 124-32.
• Whelden, C. M. 1934. Studies in the genus Fraxinus. 1. A preliminary key to winter twigs for the sections Melioides and Bumelioides. J. Arnold Arbor. 15: 118-26.
• Wilhour, R. G. 1970. The influence of ozone on white ash (Fraxinus americana L.). Ph.D. Dissertation Pennsylvania State Univ., University Park, PA, (Pennsylvania State Univ. CAES Pub. No. 188-71. 86 pp)
• Wilson, K. A.; Wood, C. E. 1959. The genera of Oleaceae in the southeastern United States. J. Arnold Arbor. 40(4): 369-84.
• Winstead, J. E.; Smith, B. J.; Wardell, G. I. 1977. Fruit weight clines in populations of ash, ironwood, cherry, dogwood, and maple. Castanea 42: 56-60.
• Woodcock, H.; Patterson, W. A.; Davies, K. M. 1993. The relationship between site factors and white ash decline in Massachusetts. Forest Ecol. & Manag. 60: 271-90.
• Wright, J. W. 1944. Genotypic variation in white ash. J. Forest. 42(7): 489-95.
• Wright, J. W. 1959. Silvical characteristics of green ash. (Upper Darby, PA)
• Wright, J. W. 1959. Silvical characteristics of white ash (Fraxinus americanum).
• Wright, J. W. 1944. Ecotypic differentiation in red ash. J. Forest. 42(8): 591-7.
• Wright, J. W. 1945. Epidermal characters in Fraxinus. Proc. Indiana Acad. Sci. 54: 84-90.
• Wright, J. W. 1944. Epidermal characters in Fraxinus. Proc. Indiana Acad. Sci. 54: 84-90.
• Wright, J. W. 1953. Pollen dispersion studies: some practical applications. J. Forest. 51: 114-8.
• Wright, J. W. 1952. Pollen dispersion of some forest trees.
• Wright, J. W. 1957. New chromosome counts in Acer and Fraxinus. Morris Arbor. Bull. 8: 33-4.
• Yetter, T. C.; Runkle, J. R. 1986. Height growth rates of canopy tree species in southern Appalachian gaps. Castanea 51: 157-67.
• Zamski, E.; Zimmermann, M. H. 1979. Sieve tube longevity in white ash (Fraxinus americana) studied with a new histochemical test for the identification of sugar. Canad. J. Bot. 57: 650-6.
• Zimmermann, M. H.; Jeje, A. A. 1981. Vessel-length distribution in stems of some American woody plants. Canad. J. Bot. 59: 1882-1892.