Genus: Alnus
Nomenclature
Alnus Mill., Gard. Dict. Abr. ed. 4., v.1 [“AL”]. 1754. LECTOTYPE: Alnus vulgaris Hill (=Alnus glutinosa (L.) Gaertn.) designated by Britton & Brown (1913).Key to the species of Alnus
1. Single-trunked tree...Alnus glutinosa1. Multi-stemmed shrub...2
2. Leaves doubly serrate, undersides not resin-coated; bark
dark grayish to reddish-brown, with prominent whitish lenticels...Alnus incana
2. Leaves simply serrate or
serrulate, undersides resin-coated; bark light gray, without prominent
lenticels...3
3. Leaves serrate, mostly ovate, apices acute...Alnus viridis
3. Leaves serrulate, mostly
obovate, apices rounded...Alnus
serrulata
List of Alnus Species
References to Alnus
- Abbe, E. C. 1933. The inter-relationship of the genera of the Betulaceae, based on anatomical studies of the inflorescence, the flowers, and the secondary xylem. MS Thesis Harvard Univ.,
- Abbe, E. C. 1935. Studies in the phylogeny of the Betulaceae. I. Floral and inflorescence anatomy and morphology. II. Extremes in the range of variation of floral and inflorescence morphology. Bot. Gaz. 97: 1-67.
- Abbe, E. C. 1930. The anatomy and morphology of the staminate inflorescence and flowers of the Betulaceae. M.S. Thesis Cornell Univ., Ithaca, NY33 plates + 30 p.
- Altpeter, L. S. 1944. Use of vegetation in control of streambank erosion in Northern New England. J. Forest. 42(2): 99-107.
- Anderson, E.; Abbe, E. C. 1934. A quantitative comparison of specific and generic differences in the Betulaceae. J. Arnold Arbor. 15: 43-9.
- Anonymous 1979. Symbiotic nitrogen fixation in actinomycete-nodulated plants. Bot. Gaz. 140(Supplement): 1-126.
- Ashburner, K. 1986. Alnus: a survey. The Plantsman 8(3): 170-88.
- Blom, J.; Roelofson, W.; Akkermans, A. D. L. 1981. Assimilation of nitrogen in root nodules of alder, Alnus glutinosa. New Phyt. 89: 321-6.
- Bond, G. 1956. Evidence for fixation of nitrogen by root nodules of alder (Alnus) under field conditions. New Phyt. 55: 147-53.
- Bond, G. 1967. Nitrogen fixation in some non-legume root nodules. Phyton 24: 57-66.
- Bond, G.; Fletche, W. W.; Ferguson, T. P. 1954. The development and function of the root nodules of Alnus, Myrica and Hippophae. Pl. & Soil 5: 309-23.
- Bormann, B. T.; Bormann, F. H.; et al. et.al. 1993. Rapid N2 fixation in pines, alder, and locust: evidence from the sandbox ecosystem study. Ecology 74: 583-98.
- Bousquet, J.; Cheliak, W. M.; Lalonde, M. 1987. Allozyme variability in natural populations of green alder (Alnus crispa) in Quebec. Genome 29: 345-52.
- Bousquet, J.; Cheliak, W. M.; Lalonde, M. 1987. Genetic diversity within and among 11 juvenile populations of green alder (Alnus crispa) in Canada. Physiol. Pl. (Copenhagen) 70: 311-8.
- Bousquet, J.; Cheliak, W. M.; Lalonde, M. 1987. Genetic differentiation among 22 native populations of green alder (Alnus crispa) in central Quebec. Canad. J. Forest Res. 17: 219-27.
- Bousquet, J.; Cheliak, W. M.; Lalonde, M. 1988. Allozyme variation within and among mature populations of speckled alder (Alnus rugosa) and relationships with green alder (Alnus crispa). Amer. J. Bot. 75(11): 1678-86.
- Briggs, D. G.; DeBell, D. S.; Atkinson, W. A. (eds.) (1978): 1978. Utilization and management of alder. Portland.
- Britton, Nathaniel L. 1904. An undescribed species of Alnus (A. noveboracensis). Torreya 4: 124.
- Brown, D. M. 1941. The vegetation on Roan Mountain, a phytosociological and successional study. Ecol. Monogr. 11: 61-97.
- Bunce, J. A.; Miller, L. N. 1976. Differential effects of water stress on respiration in the light in woody plants from wet and dry habitats. Canad. J. Bot. 54(21): 2457-64.
- Bunce, J. A.; Miller, L. N.; Chabot, B. F. 1977. Competative exploitation of soil water by five eastern North American tree species. Bot. Gaz. 138(2): 168-73.
- Carpenter, S. B.; Smith, N. D. 1975. Stomatal distribution and size in southern Appalachian hardwoods. Canad. J. Bot. 53: 1153-6.
- Carpenter, S. B.; Smith, N. D. 1979. Variation in shade leaf thickness among urban trees growing in metropolitan Lexington, Kentucky. Castanea 44: 94-8.
- Carpenter, S. B.; Smith, N. D. 1981. A comparative study of leaf thickness among southern Appalachian hardwoods. Canad. J. Bot. 59: 1393-6.
- Chamberlain, C. 1898. Winter characters of certain sporangia. Bot. Gaz. 25: 124-8.
- Chen, Z. D.; Manchester, S. R.; Sun, H. Y. 1999. Phylogeny and evolution of the Betulaceae as inferred from DNA sequences, morphology, and paleobotany. Amer. J. Bot. 86: 1168-1181.
- Chen, Z.; Li, J. 2004. Phylogenetics and biogeography of Alnus (Betulaceae) inferred from sequences of nuclear ribosomal DNA ITS region. Int. J. Plant Sci. 165: 325-335.
- Clarkson, R. B. 1960. Notes on the distribution of Alnus crispa in eastern North America. Castanea 25: 83-6.
- Dalton, D. A.; Naylor, A. W. 1975. Studies on nitrogen fixation by Alnus crispa. Amer. J. Bot. 62(1): 76-80.
- Daly, G. T. 1966. Nitrogen fixation by nodulated Alnus rugosa. Canad. J. Bot. 44: 1607-21.
- Duncan, W. H. 1950. Preliminary reports on the flora of Georgia. 2. Distribution of 87 trees. Amer. Midl. Naturalist 43: 742-61.
- Echternach, J. L.; Rose, R. K. 1987. Use of woody vegetation by beavers in southeastern Virginia USA. Virginia J. Sci. 38: 226-232.
- Ehrenfeld, J. G.; Gulick, M. 1981. Structure and dynamics of hardwood swamps in the New Jersey Pine Barrens: contrasting patterns in trees and shrubs. Amer. J. Bot. 68: 471-81.
- Ferguson, T. P.; Bond, G. 1953. Observations on the formation and function of the root nodules on Alnus glutinosa. Ann. Bot. 17: 175-88.
- Fernald, M. L. 1945. Some North American Corylaceae (Betulaceae). II. Eastern North American representatives of Alnus incana. Rhodora 47(563): 333-61.
- Fernald, M. L. 1904. The green alders of New England. Rhodora 6: 162-3.
- Fremstad, E. 1983. Role of black alder (Alnus glutinosa) in vegetation dynamics in west Norway. Nordic J. Bot. 3: 393-410.
- Furlow, J. J. 1979. The systematics of the American species of Alnus (Betulaceae). Rhodora 81: 1-121, 151-248.
- Furlow, J. J. 1976. Nomenclatural changes in (Alnus) (Betulaceae). Ann. Missouri Bot. Gard. 63: 380, 381.
- Furlow, J. J. 1983. Evolutionary divergence and classification of the Alnus viridis complex (Betulaceae). Amer. J. Bot. 70(Suppl.): 114.
- Furlow, J. J. 1990. The genera of Betulaceae in the southeastern United States. J. Arnold Arbor. 71: 1-67.
- Furlow, J. J. 1982. The alders of North America. Int. Dendrol. Soc. Year Book 1981: 115-9.
- Furlow, J. J. 1981. The Alnus incana complex in North America. Ohio J. Science 81 (Apr. Prog. Abstr.): 33. (Ohio Acad. Sci., 90th Annual Meeting April 24-26, 1981, College of Wooster.)
- Genys, J. B. 1988. Intraspecific variation among 28 different sources of black alder, Alnus glutinosa (Betulaceae). Castanea 53: 71-9.
- Gibson, J. P. et.al. 2008. Comparison of population genetic diversity between a rare, narrowly distributed species and a common, widespread species of Alnus (Betulaceae). Amer. J. Bot. 95: 588-596.
- Green, W. E. 1947. Effect of water impoundment on tree mortality and growth. J. Forest. 45: 118-20.
- Hacke, U.; Sauter, J. J. 1996. Drought-induced xylem dysfunction in petioles, branches, and roots of Populus balsamifera L. and Alnus glutinosa (L.) Gaertn. Pl. Physiol. (Rockville) 111: 413-417.
- Hall, J. W. 1952. The comparative anatomy and phylogeny of the Betulaceae. Bot. Gaz. 113(3): 235-70.
- Hardin, J. W. 1952. The Juglandaceae and Corylaceae of Tennessee. Castanea 17: 78-89.
- Hardin, J. W.; Bell, J. M. 1986. Atlas of foliar surface features in woody plants: IX. Betulaceae of eastern United States. Brittonia 38(2): 133-44.
- Hawker, L. E.; Fraymouth, J. 1951. A re-investigation of the root-nodules of species of Elaeagnus, Hippophaë, Alnus, and Myrica, with special reference to the morphology and life histories of the causative organisms. J. Gen. Microbiol. 5: 369-86.
- Herron, P.M. et.al. 2007. Invasive plants and their ecological strategies: prediction and explanation of woody plant invasion in New England. Diversity and Distributions 13: 633-644.
- Hollick, C. A. 1922. Alnus rugosa. Addisonia 7: 25-6.
- Huenneke, L. F. 1985. Spatial distribution of genetic individuals in thickets of Alnus incana ssp. rugosa, a clonal shrub. Amer. J. Bot. 72: 152-8.
- Huenneke, L. F. 1987. Demography of a clonal shrub, Alnus incana ssp. rugosa (Betulaceae). Amer. Midl. Naturalist 117: 43-55.
- Huenneke, L. F.; Marks, P. L. 1987. Stem dynamics of the shrub Alnus incana ssp. rugosa: transition matrix models. Ecology 68: 1234-42.
- Hurd, T. M.; Schwintzer, C. R. 1996. Formation of cluster roots in Alnus incana ssp. rugosa and other Alnus species. Canad. J. Bot. 74(11): 1684-6.
- Hylander, N. 1957. On cut-leaf and small leaved forms of Alnus glutinosa and Alnus incana. Svensk Bot. Tidskr. 51: 437-53.
- Jobidon, R.; Thibault, J. R. 1981. Allelopathic effects of balsam poplar on green alder germination. Bull. Torrey Bot. Club 108: 413-8.
- Khramova, E. L. 1996. Palynomorphology of the genus Alnus (Betulaceae). Bot. Zhurn. (Moscow & Lenengrad) 81: 42-53. (In Russian; English summary)
- Kikuzawa, K. 1980. Why do alder leaves fall in summer? Jap. J. Ecol. 30: 359-68.
- Koike, T. 1995. Effects of CO2 in interaction with temperature and soil fertility on the foliar phenology of alder, birch, and maple seedlings. Canad. J. Bot. 73(2): 149-57.
- Lavoie, C.; Jean, M.; Bouchard, A. 1991. Habitat su Sumac a vernis, Rhus vernix, a sa limite nord de distribution (Haut-Saint-Laurent, Quebec). Canad. Field-Naturalist 105: 71-7. (In French, English summary. Also Acer & Alnus)
- Love, D. 1968. Nomenclatural notes on Mt. Washington plants. Taxon 17: 89.
- Lovell, J. H. 1897. Fertilization of Alnus incana and Salix discolor. Bull. Torrey Bot. Club 24: 264-5.
- Lowry, G. L.; Brokaw, F. C.; Breeding, C. H. J. 1962. Alder for restoring coal spoils in Ohio. J. Forest. 60(3): 196-9. (A. glutinosa & A. incana)
- Lumis, G. P.; Hofstra, G.; Hall, R. 1973. Sensitivity of roadside trees and shrubs to aerial drift of deicing salt. Hortscience 8: 475-7.
- Lumis, G. P.; Hofstra, G.; Hall, R. 1975. Salt damage to roadside plants. J. Arboric. 1(1): 14-6.
- Mackintosh, A.; Bond, G. 1970. Diversity in the nodular endophytes in Alnus and Myrica. Phyton (Buenos Aires) 27: 79-90.
- Malloch, D.; Malloch, B. 1981. The mycorrhizal status of boreal plants: species from northeastern Ontario. Canad. J. Bot. 59: 2167-72.
- Matlack, G. R. 1987. Diaspore size, shape, and fall behavior in wind-dispersed plant species. Amer. J. Bot. 74(8): 1150-60.
- Matthews, R. F. 1992. Alnus viridis ssp. crispus. ()
- McDermott, R. E. 1954. Effects of saturated soil on seedling growth of some bottomland hardwood species. Ecology 35: 36-41.
- McDermott, R. E. 1953. Light as a factor in the germination of some bottomland hardwood species. J. Forest. 51: 203-4.
- McIninch, S. M.; Garbisch, E. W.; Biggs, D. 1994. The benefits of wet-acclimating woody wetland plant species. Wetland Journal 6: 19-23.
- McInteer, B. B. 1947. Soil preference of some plants as seen in Kentucky. Castanea 12: 1-8.
- McVean, D. N. 1953. Regional variation of Alnus glutinosa (L.) Gaertn. in Britain. Watsonia 3: 26-32.
- McVean, D. N. 1955. Ecology of Alnus glutinosa (L.) Gaertn. II. Seed distribution and germination. J. Ecol. 43: 61-71.
- McVean, D. N. 1956. Ecology of Alnus glutinosa (L.) Gaertn. III. Seedling establishment. J. Ecol. 44: 195-218.
- McVean, D. N. 1956. Ecology of Alnus glutinosa (L.) Gaertn. IV. Root system. J. Ecol. 44: 219-25.
- Meurer-Grimes, B. 1995. New evidence for the systematic significance of acylated spermidines and flavonoids in pollen of higher Hamamelidae. Brittonia 47(2): 130-42.
- Mian, S.; Bond, G. 1978. The onset of nitrogen fixation in young alder plants and its relation to differentiation in the nodular endophyte. New Phyt. 80: 187-92.
- Mitchell, C. C.; Niering, William A. 1993. Vegetation change in a topogenic bog following beaver flooding. Bull. Torrey Bot. Club 120(2): 136-47.
- Mitchell, R. S. (eds.) (1990): 1990. Betulaceae through Cactaceae of New York State. Vol. NY State Museum Bull. 476. Univ. of the state of NY, Albany, NY. , 93 pages.
- Molina, R. 1981. Ectomycorrhizal specificity in the genus Alnus. Canad. J. Bot. 59: 325-34.
- Murphy, J. F.; Miller, O. K. 1994. Mycorrhizal synthesis with Alnus serrulata (Ait.) Willd. Castanea 59: 156-66.
- Paratley, R. D. 1986. Vegetation-environment relations in a conifer swamp in central New York. J. Torrey Bot. Soc. 113: 357-371.
- Rich, A. B. 1889. Heterogamy in Alnus serrulata. Bull. Torrey Bot. Club 16: 112.
- Ricklefs, R. E.; Matthew, K. K. 1982. Chemical characteristics of the foliage of some deciduous trees in southeastern Ontario. Canad. J. Bot. 60: 2037-45.
- Rose, S. L. 1980. Mycorrhizal associations of some actinomycete nodulated nitrogen-fixing plants. Canad. J. Bot. 58: 1449-54.
- Schalin, I. 1968. Germination analysis of grey alder (Alnus glutinosa) seeds. In: Biology of Alder. Northwest Sci. Assoc. 40th Annual meeting Symp. Proc. Pacific Northwest Forest and Range Exp. Sta., Portland, OR. , 107-13 pages.
- Schrader, J. A. et.al. 2006. Differences in shade tolerance help explain varying success of two sympatric Alnus species. Int. J. Plant Sci. 167: 979-989.
- Schwintzer, C. R.; Tjepkema, J. D. 1997. Field nodules of Alnus incana ssp. rugosa and Myrica gale exhibit pronounced acetylene-induced declines in nitrogenase activity. Canad. J. Bot. 75: 1415-23.
- Sequerra, J. et al. et.al. 1994. Root hair infection process and myconodule formation on Alnus incana by Penicillium nodositatum. Canad. J. Bot. 72(7): 955-62.
- Silvester, W. B.; Silvester, J. K.; Torrey, J. G. 1988. Adaptation of nitrogenase to varying oxygen tension and the role of the vesicle in root nodules of Alnus incana ssp. rugosa. Canad. J. Bot. 66: 1772-9.
- Steele, F. L. 1961. Introgression of Alnus serrulata and Alnus rugosa. Rhodora 63: 297-304.
- Stibolt, V. M. 1981. The distribution of Alnus maritima Muhl. ex Nutt. (Betulaceae). Castanea 46: 195-200.
- Suringar, J. V. 1931. Nomenklaturalia. Mitt. Deutsch. Dendrol. Ges. 43: 199-217. (In German)
- Tarrant, R. F. 1968. Some effects of alder on the forest environment. In: Biology of Alder. Portland. , 193 pages.
- Trappe, J. M.; Franklin, J. F.; Tarrant, R. F.; Hansen, G. M. (eds.) (1968): 1968. Biology of Alder; Proceedings of a Symposium held at Northwest Scientific Association Fortieth Annual Meeting, Pullman, Washington, April 14-15, 1967. U.S. Pacific Northwest Forest and Range Exp. Sta., Portland. , 292 pages.
- Turrill, W. B. 1962. Alnus viridis ssp. crispa. Bot. Mag. 173: 382.
- Vaartaja, O. 1954. Photoperiodic ecotypes of trees. Canad. J. Bot. 32: 392-9.
- Van Deelen, T. R. 1991. Alnus rugosa. ()
- Villamil, C. B.; Fairbrothers, D. E. 1974. Comparative protein population investigation of the Alnus serrulata-rugosa complex. Biochem. Syst. Ecol. 2: 15-20.
- Wherry, Edgar T. 1960. Intermediate occurrences of Alnus crispa. Castanea 25: 135.
- Wolpert, J. 1910. Vergleichende Anatomie und Entwicklungsgeschichte von Alnus, Alnobetula und Betula. Flora 100: 37-67. (In German)
- Woodworth, R. H. 1931. Polyploidy in Betulaceae. J. Arnold Arbor. 12: 206-17.
- Woodworth, R. H. 1929. Cytological studies in the Betulaceae. II. Corylus and Alnus. Bot. Gaz. 88: 383-99.
- Woodworth, R. H. 1930. Cytological studies in the Betulaceae. III. Parthenogenesis and polyembryony in Alnus rugosa. Bot. Gaz. 89: 402-9.
- Woodworth, R. H. 1929. Parthenogenesis and polyploidy in Alnus rugosa (Du Roi) Spreng. Science 70: 192-3.
- de Poucques, M. L. 1949. Etudes caryologiques sur les Fagales. I. Le genre Alnus. Bull. Mus. Hist. Nat. (Paris) 21(2): 147-52.